Genus Horribates Muma 1962

Horribates Muma, 1962: 7; Muma, 1970a: 29; Muma, 1976: 18; Muma, 1989: 15; Harvey 2003e: 248; Brookhart and Brookhart 2006: 301 (key), 314:.


    Right chelicera, pedipalp, and genital opercula of female Horribates spinigerus Muma.


Type species:

Horribates spinigerus Muma 1951


Diagnosis:

Eremobatid solifuges with robust, movable spines on the pedipalps.


Original description:

Muma 1962: 7:

7 - HORRIBATES, NEW GENUS

Although no comparative diagnosis of this genus can be made in the absence of males, there can be no question concerning the unique generic status of the generotype described below. The long slender chelicerae, the extremely short palpal tarsus, and the unusual spination of the palpus are distinctive within the Eremobatidae. Only one tiny aborted tarsal claw could be found on the first leg, so the genus is tentatively placed in the Eremobatinae.

GENEROTYPE: Horribates spinigerus, new species.


Subsequent accounts:

Muma 1970a 3 (key), 29:

3 -KEY TO SUBFAMILIES AND GENERA OF EREMOBATIDAE
(MALES AND FEMALES)

1. Leg I with 1 claw; large, robust, short legged species ...................... Eremobatinae Roewer-2

    Leg I with 2 claws; small, slender, long legged species ....................... Therobatinae Muma-5

2. Palpus with 2 ventral rows of long, robust, spinelike, erectile and deflectile setae on femur,
   tibia, metatarsus, and tarsus ...................................................................... Horribates Muma

    Palpus with or without spinelike setae; when present, apparently not movable and not on
   metatarsus and tarsus .......................................................................................................... 3

(MALES ONLY)

3. Fixed finger short and dorsally lobate or sculptured ................................ Eremothera Muma

    Fixed finger elongate and need1elike or stylelike ................................................................ 4

4. Mesoventral groove of fixed finger extending to base of finger; apical seta of flagellum-
  complex flattened and plumose ................................................................ Eremobates Banks

    Mesoventral groove of fixed finger not extending to base of finger; apical seta of flagellum-
  complex tubular, at most striate ............................................................... Eremorhax Roewer

5. Dorsal setae of flagellum complex simple and tubular ........................... Eremochelis Roewer

    Dorsal setae of flagellum-complex striate, plumose, spatulate, or otherwise modified .......
   ............................................................................................................................................... 6

6. Fixed finger distinctly sinuate, bent upward and bent or curved downward; dorsal setae
  plumose ......................................................................................................... Chanbria Muma

29. - Genus Horribates Muma
Horribates Muma, 1962, p. 7.


Muma 1976: 18

18 -    Genus Horribates Muma, 1962

To date, this is a monotypic genus based on female characters. Males are presently unknown.


Muma 1989: 3 (key), 18:

    3 -     Key to Subfamilies and Genera of Eremobatidae

  (Males Only)

1. Leg 1 with 1 claw; chelicerae about twice as long as wide; small to large species ...............
   ........................................................................................................Eremobatinae Roewer-2

1'. Leg 1 with 2 claws; chelicerae 2.5–3 times longer than wide; tiny to moderate sized
   species ......................................................................................................... Therobatinae-6

2. Fixed cheliceral finger long, style-like or needle-like; mesoventral groove a crease, slot or
   cup-like structure; moderate-sized to large species .......................................................... 3

2'. Fixed cheliceral finger short, sculptured and flanged; mesoventral groove a trough-like
    slot; moderate sized species ................................................................ Eremothera Muma

3. Mesoventral groove an indistinct hollow or crease that does not extend to the base of
    the fixed finger; movable finger dentition greatly reduced ................................................
   ....................................................................................... Arenotherus Brookhart and Muma

3'. Mesoventral groove a distinct crease, cup, or slot that may or may not extend to base of
   fixed finger; movable finger dentition normal or increased .............................................. 4

4. Mesoventral groove short, not extending to base of fixed finger; apical striate or
   plumose setae of male flagellum complex not obviously modified or flattened .............. 5

4'. Mesoventral groove long, extending to base of fixed finger; apical plumose seta of male
   flagellum complex obviously enlarged or flattened covering part of the mesoventral
   groove..................................................................................................... Eremobates Banks

5'. Palpal metatarsus, tibia, and femur provided with 2 ventral rows of enlarged, robust,
    spine-like, obviously movable setae ....................................................... Horribates Muma

6. Fixed cheliceral finger strongly recurved, sigmoid, or S-shaped; mesoventral groove
    absent; both dorsal and ventral flagellum complex setae plumose, moderate sized
    species........................................................................................................Chanbria Muma

6'. Fixed cheliceral finger style-like or needle-like, straight, curved, or undulate but not
    S-shaped or sigmoid; mesoventral groove present; dorsal flagellum complex setae
    striate, ventral setae striate or plumose ......................................................................... 7

7. Fixed cheliceral finger with a mesoventral groove that varies from one or more indistinct
    creases to an elongate hollow or cup; small to moderate sized species ............................
   ............................................................................................................. Eremochelis Roewer

7'. Fixed cheliceral finger without a mesoventral groove; tiny to small species ......................
    ............................................................................................................ Hemerotrecha Banks

18 - Genus Horribates Muma, 1962

Horribates Muma, 1962, p. 7; Muma, 1970, p . 29. Females only .

Collection of additional specimens of type species and 2 undescribed species has greatly increased our knowledge of this genus.

Key to species of genus Horribates

(Females)

  1.  Species with movable, spine-like setae on palpal tarsi; females 20 mm. or more in length;
      opercular pits directed ventrally and medially located ......................... H. spinigerus Muma

  1'. Species with no movable, spine-like setae on palpal tarsi; females 15 mm. or less in length;
      opercular pits not directed ventrally and not located medially .......................................... 2

  2.  Females with 5 pairs of movable , spine-like setae on palpal femora; opercular pits
      relatively large , located laterally and directed laterally ............................... H. bantai n. sp .

  2'. Females with 3 pairs of movable, spine-like setae on palpal femora; opercular pits small,
      located laterally and directed ventrolaterally ............................................. H. minimus n. sp.

 

Harvey 2003e: 248:

248 - Genus Horribates Muma

Horribates Muma, 1962: 7; Muma, 1970a: 29; Muma, 1976: 18; Muma, 1989: 15.

Type species: Horribates spinigerus Muma, 1962, by original designation.

Distribution: south-western U.S.A.


Brookhart and Brookhart 2006: 301 (key), 314:

301 -    Key to Subfamilies and Genera of Eremobatidae (males only)

(Taken from Muma 1987)

1. Leg 1 with 1 claw; chelicerae about twice as long as wide; small to large species ...............
   ........................................................................................................Eremobatinae Roewer-2

     Leg 1 with 2 claws; chelicerae 2.5–3 times longer than wide; tiny to moderate sized
   species ......................................................................................................... Therobatinae-6

2. Fixed cheliceral finger long, style-like or needle-like; mesoventral groove a crease, slot or
   cup-like structure; moderate-sized to large species .......................................................... 3

    Fixed cheliceral finger short, sculptured and flanged; mesoventral groove a trough-like
    slot; moderate sized species ........................................................................... Eremothera

3. Mesoventral groove an indistinct hollow or crease that does not extend to the base of
    the fixed finger; anterior tooth absent ............................................................. Eremorhax

    Mesoventral groove a distinct crease, cup, or slot that may or may not extend to base of
   fixed finger; anterior tooth present ................................................................................... 4

4. Mesoventral groove short, not extending to base of fixed finger; apical striate or
   plumose setae of male flagellum complex not obviously modified or flattened .............. 5

    Mesoventral groove long, extending to base of fixed finger; apical plumose seta of male
   flagellum complex obviously enlarged or flattened; covering part of the mesoventral
   groove................................................................................................................ Eremobates

5. Palpal metatarsus, tibia, and femur provided with enlarged spine-like seta, but not
   robust, not aligned in a ventral row, not movable ........................................... Eremocosta

    Palpal metatarsus, tibia, and femur provided ventrally with robust, obviously movable
   spinelike setae .................................................................................................... Horribates

6. Fixed cheliceral finger strongly recurved, sigmoid, or S-shaped; mesoventral groove
    absent; both dorsal and ventral flagellum complex setae plumose, moderate sized
    species...................................................................................................................Chanbria

    Fixed cheliceral finger style-like or needle-like, straight, curved, or undulate but not
   S-shaped or sigmoid; mesoventral groove present; dorsal flagellum complex setae
   striate, ventral setae striate or plumose ........................................................................... 7

7. Fixed cheliceral finger with a mesoventral groove that may vary from a distinct crease to
   an elongate hollow or cup; dorsal flagellum complex simple or tubular; small to
   moderate sized species .................................................................................... Eremochelis

   Fixed cheliceral finger without a mesoventral groove; dorsal flagellum complex hooked or
   spatulate; tiny to small species ..................................................................... Hemerotrecha

314 -   Genus Horribates Muma 1962

Horribates Muma 1962:7.


Notes:


Included species



Bibliography: