Genus Eremochelis Roewer 1934

Eremochelis Roewer, 1934: 556 (figs. 320d, n), 570; Muma, 1970, p. 30; Harvey, 2003e: 249; Brookhart and Brookhart 2006: 301 (key) 315.

Therobates Muma, 1951, p. 85; Muma, 1962, p. 9; Muma 1989: 3 (key), 18.



Type species:

Eremochelis insignitus (Roewer 1934)


Diagnosis:

From Muma (1951): "Large- to small-sized Therobatinae. Fixed finger of male chelicera with a mesal or mesoventral groove. Flagellum complex composed of a dorsal row or group of simple tubular bristles, a mesal row or group of plumose bristles, and a ventral or basal row or group of simple tubular bristles. The plumose bristles partially or completely cover the mesoventral groove. First postspiracular abdominal sternite of males with ctenidia on its dorsal margin. Mesal tooth of movable finger of males and females present or absent. Opercula of females variable."



Original description:

Roewer 1934: 570:

570 - "8. Gen. Eremochelis nov. gen.

Genotypus: E. insignitus nov. spec.

Eremobatinae, deren 2. und 3. Tarsus ventral mit jeweils 2.2.2.4.6 Dornen und deren 4. Tarsus ventral mit 2.2.2.2.2.2/2/1.2.6 Dornen bewehrt ist (Abb. 320 d, n).

Nur 1 Art:

1. Eremochelis insignitus nov. spec.

♂: Flagellum-Komplex aus einem Büschel einfacher Borsten bestehend; unbeweglicher Finger gerade, ohne Basalbuckel, seine Schneide etwas geschweift, mit 6 lateralen (der 1. am größten) und 3 medialen (der 2. am größten) Wangenzähnen; beweglicher Finger mit 1 Vorder-, 1 Zwischen-, 1 Hauptzahn, ohne 1 medialen Wangenzahn! Gebiß (Abb. 322, k); Pedipalpen nur behaart, Tibia und Metatarsus dorsal und ventral mit Zylinderborsten, Metatarsus ventral ohne Scopula, apikal auffällig dicker als basal; 2. Tibia dorsalapikal mit 1 und 3. Tibia hier mit 2 Dornen; 2. und 3. Metatarsus dorsal mit 6 Dornen der Haupt- und 2 Dornen der Nebenreihe, Tarsalkrallen auffällig lang und dünn; 5. Sternit des Opisthosoma mit 2: 2 Ctenidien (deren Form Abb. 325, c); Färbung vollig blaßgelb, nur Metatarsus und Tarsus der Pedipalpen dunkelbraun, Malleoli weiß; Körperlänge 10 mm. - ♀: Unbeweglicher Finger mit 4 Vorderzähnen, davor auf der Schneide eine Reihe aus 5 Sägezähnchen, dahinter 1 Zwischen-, 1 Haupt-, 4 laterale und 3 mediale Wangenzähne; beweglicher Finger mit 1 Vorder-, 1 Zwischen- und 1 Hauptzahn, ohne 1 medialen Wangenzahn, doch vor dem Vorderzahn auf der Schneide mit 8 Sägezähnchen besetzt; Gebiß (Abb. 327, d); Pedipalpen wie beim ♂ beborstet, doch ohne Zylinderborsten, Metatarsus wie beim ♂ geformt; Bedornung der Beine wie beim ♂; 5. Sternit ohne Ctenidien; Sternithälften des Genitalsegmentes (Abb. 324, r); Färbung wie beim ♂; Körperlänge 11 mm.

♂, ♀ - Californien (genauer Fundort ?) - (vidi 1 ♂, ♀, Typus).


Subsequent accounts:

 

Muma 1951 (as Therobates): 85:

85 - "KEY TO GENERA

MALES

    1. Dorsal bristles of flagellum complex simple and tubular........................... Therobates

       Dorsal bristles of flagellum complex plumose, striate, or otherwise modified ..........2

    2. Fixed finger distinctly sinuate ........................................................................ Chanbria

       Fixed finger straight or at most weakly curved or undulate .................. Hemerotrecha

THEROBATES1, NEW GENUS

Eremobates of authors, in part.

Large- to small-sized Therobatinae. Fixed finger of male chelicera with a mesal or mesoventral groove. Flagellum complex composed of a dorsal row or group of simple tubular bristles, a mesal row or group of plumose bristles, and a ventral or basal row or group of simple tubular bristles. The plumose bristles partially or completely cover the mesoventral groove. First postspiracular abdominal sternite of males with ctenidia on its dorsal margin. Mesal tooth of movable finger of males and females present or absent. Opercula of females variable.

GENOTYPE: Therobates bilobatus, new species.

KEY TO GROUPS

MALES

    1. Apical plumose bristle of flagellum complex conspicuously enlarged and flattened....
       ................................................................................................................ branchi group

        Apical plumose bristle of flagellum complex not conspicuously enlarged ................. 2

    2. Mesoventral groove of fixed finger distinct, movable finger not modified ..................
       .............................................................................................................. bilobatus group

        Mesoventral groove of fixed finger indistinct, movable finger modified apically .........
     ............................................................................................................. imperialis group

1 Greek, ther, a wild animal, plus bates (combining form), a climber.

 

Muma 1962 (as Therobates): 9:

9 - The key below is a revision of that on page 85 in Muma (1951), to include the new arcus group, andreasana group, and striodorsalis group, described below.

KEY TO GROUPS

MALES

    1.  Apical plumose bristle of flagellum complex conspicuously enlarged and flattened..........2

          Apical plumose bristle of flagellum complex not conspicuously enlarged and flattened...4

    2.  Groove of fixed finger mesoventral in position .................................................................. 3

          Groove of fixed finger dorsal to dorsomesial in position ...................... striodorsalis group

    3.  Mesoventral groove composed of weak ridges and hollows; movable finger modified
         apically .................................................................................................. andreasana group

         Mesoventral groove a distinct hollow cup; movable finger not modified apically..............
        ..................................................................................................................... branchi group

    4.  Mesoventral groove distinct; movable finger not modified apically................................ 5

         Mesoventral groove indistinct; movable finger modified apically............imperialis group

    5.  Mesoventral groove a narrow slot without distinct carinae .......................... arcus group

         Mesoventral groove a wide hollow cup with distinct carinae ................. bilobatus group

 

 

Muma 1970a: 3 (key), 30:

   3 - Key to Subfamilies and Genera of Eremobatidae

(MALES AND FEMALES)

    1.  Leg I with 1 claw; large, robust, short legged species .............Eremobatinae Roewer - 2

         Leg I with 2 claws; small, slender, long legged species ................Therobatinae Muma-5

    2.  Palpus with 2 ventral rows of long, robust, spinelike, erectile and deflectile setae on
        femur, tibia, metatarsus, and tarsus ...................................................Horribates Muma

         Palpus with or without spinelike setae; when present, apparently not movable
        and not on metatarsus and tarsus ................................................................................ 3

(MALES ONLY)

    3.  Fixed finger short and dorsally lobate or sculptured .................... Eremothera Muma

         Fixed finger elongate and needlelike or stylelike ....................................................... 4

    4.  Mesoventral groove of fixed finger extending to base of finger; apical seta of
        flagellum-complex flattened and plumose .................................... Eremobates Banks

         Mesoventral groove of fixed finger not extending to base of finger; apical seta of
        flagellum-complex tubular, at most striate....................................Eremorhax Roewer

    5.  Dorsal setae of flagellum complex simple and tubular.................Eremochelis Roewer

         Dorsal setae of flagellum-complex striate, plumose, spatulate, or otherwise
        modified ....................................................................................................................6

    6.  Fixed finger distinctly sinuate, bent upward and bent or curved downward; dorsal
        setae plumose.....................................................................................Chanbria Muma

         Fixed finger straight, weakly curved, undulate or bent downward at tip; dorsal setae
        striate, spatulate or hooked .......................................................Hemerotrecha Banks

    30 -  Genus Eremochelis Roewer

   Eremochelis Roewer, 1934, p. 570.

   Therobates Muma, 1951, p. 85 (new synonymy).

    KEY TO SPECIES-GROUPS

    (MALES)

    1.  Apical plumose bristle of the flagellum complex conspicuously enlarged and
        flattened......................................................................................................................2

         Apical plumose bristle of flagellum complex not enlarged or flattened......................4

    2.  Groove of fixed finger dorsal to dorsomesial in position................striodorsalis group

          Groove of fixed finger mesoventral in position .........................................................3

    3.  Mesoventral groove weakly hollowed and ridged; movable finger modified
        apically.............................................................................................andreasana group

        Mesoventral groove a distinct cup or slot; movable finger not modified apically
       .................................................................................................................branchi group

    4.  Mesoventral groove indistinct; movable finger; movable finger modifed
        apically.................................................................................................imperialis group

        Mesoventral groove distinct; movable finger not modified apically............................5

    5.  Mesoventral groove a wide hollow cup with distinct carinae..............bilobatus group

        Mesoventral groove a narrow slot without distinct carinae.......................arcus group


Muma 1989: 3 (key), 18-19:

   3 -  Key to Subfamilies and Genera of Eremobatidae

(Males Only)

    1.  Leg 1 with one claw; chelicerae about twice as long as wide; small to large
       species..............................................................................................Eremobatinae....2

    1'. Leg 1 with 2 claws; chelicerae 2.5 to 3 times longer than wide; tiny to moderate-sized
        species..............................................................................................Therobatinae....6

    -----

    6.  Fixed chelicera! finger strongly recurved, sigmoid or S-shaped; mesoventral groove
       absent; both dorsal and ventral flagellum complex setae plumose; moderate-sized
       species .................................................................................................Chanbria Muma

    6'. Fixed chelicera! finger style-like or needle-like and straight, curved or undulate but not
        S-shaped; mesoventral groove present or absent; dorsal flagellum complex setae striate,
        ventral setae striate or plumose.................................................................................7

    7.  Fixed cheliceral finger with a mesoventral groove that varies from one or more
        indistinct creases to an elongate hollow or cup; small to moderate-sized species
        ............................................................................................................Eremochelis Roewer

18-19 -   Genus Eremochelis Roewer

 Eremochelis Roewer, 1934, p. 570; Muma, 1970, p. 30.

 Therobates Muma, 1951, p. 85; Muma, 1962, p. 9.

 Small to large Therobatinae. Male fixed cheliceral finger with dorsomesal, mesal, or mesoventral groove. Dorsal setae of flagellum complex weakly to strongly striate, ventral setae weakly to strongly plumose. Plumose setae partially to completely cover mesal, mesoventral, or dorsomesal groove. Apical striate and apical plumose setae not especially modified on most species groups. However, in the branchi group the apical plumose seta is distinctly flattened. In the andreasana group both apical setae are strong and striate. The bilobatus species has both dorsal and ventral setae of complex plumose . Males, in addition to usual palpal clothing, often have femora, tibiae, and metatarsi provided below with one or two unequal rows of elongate spinelike setae. Female opercula extremely variable within genus but relatively consistent within species groups.

18-19 - Key to species groups of Eremochelis Roewer

  (Males)

   1. Fixed cheliceral finger with a distinct, elongate, mesoventral or dorsomesal cavity or
      slot occupying at least half of finger length.......................................................................2

   1'. Fixed chelicera! finger with short mesal creases, hollows, grooves or very short apical
      cups....................................................................................................................................4

   2. Apical plumose seta of flagellum complex distinctly to conspicuously flattened and
      widened .............................................................................................................................3

   2'. Apical plumose seta of flagellum complex obscurely or not widened and flattened
      ......................................................................................................................bilobatus group

   3. Fixed chelicera! finger without dorsal flange and with groove mesoventral in
      position...........................................................................................................branchi group

   3'. Fixed finger with dorsal flange and with groove dorsomesal in position.............................
     .................................................................................................................striodorsalis group

  4. Fixed finger straight or evenly curved ventrally; mesal groove one or more creases;
      flagellum complex apically with striate and plumose setae; movable finger with an
      apical cup or hollow....................................................................................imperialis group

  4'. Fixed finger undulate or notched ventrally; mesal groove a distinct to indistinct cup-like
      hollow; flagellum complex apically with only striate setae; movable finger with or
       without a flange .................................................................................... andreasana group

   ( Females )

  1.  Opercula provided posteriorly with a distinct mesal notch................................................2

  1'. Opercula flared but not distinctly notched posteriorly.......................................................3

  2.  Opercula commonly provided with anteriolateral pits and often with adjacent distinct
     mesal lobes at the anterior end of the notch..............................................bilobatus group

  2'. Opercula not usually provided with anteriolateral pits but often strongly constricted
       laterally and with small, triangular elevations near the posterior end of the notch..........
      ......................................................................................................................branchi group

  3.  Lateral pits of opercula indistinct, often hidden behind the lateral wings..........................
      ...............................................................................................................andreasana group

  3'. Lateral pits of opercula distinct, often invading lateral wings....................imperialis group

  Females of striodorsalis group unknown.


Harvey, 2003e: 249:

  249 - Genus Eremochelis Roewer

Eremochelis Roewer, 1934: 570; Muma, 1970a: 30; Muma, 1989: 18.

Therobates Muma, 1951: 85; Muma, 1976: 18 (synonymised by Muma, 1970a: 30).

Type locality: of Eremochelis: Eremochelis insignatis Roewer, 1934, by original designation.

of Therobates: Therobates bilobatus Muma, 1951, by original designation.

Distribution: Mexico, south-western U.S.A.


Brookhart and Brookhart 2006: 301 (key), 315:

301 -    Key to Subfamilies and Genera of Eremobatidae (males only)

(Taken from Muma 1987)

1. Leg I with 1 claw; chelicerae about twice as long as wide; small to large species .....................
   .............................................................................................................Eremobatinae Roewer-2

   Leg 1 with 2 claws; chelicerae 2.5–3 times longer than wide; tiny to moderate sized species ...
   ........................................................................................................................... Therobatinae-6

2. Fixed cheliceral finger long, style-like or needle-like; mesoventral groove a crease, slot or
   cup-like structure; moderate-sized to large species ................................................................ 3

    Fixed cheliceral finger short, sculptured and flanged; mesoventral groove a trough-like slot;
   moderate sized species .......................................................................................... Eremothera

3. Mesoventral groove an indistinct hollow or crease that does not extend to the base of the
   fixed finger; anterior tooth absent .......................................................................... Eremorhax

    Mesoventral groove a distinct crease, cup, or slot that may or may not extend to base of
   fixed finger; anterior tooth present ......................................................................................... 4

4. Mesoventral groove short, not extending to base of fixed finger; apical striate or plumose
   setae of male flagellum complex not obviously modified or flattened ................................... 5

    Mesoventral groove long, extending to base of fixed finger; apical plumose seta of male
   flagellum complex obviously enlarged or flattened; covering part of the mesoventral groove
   ................................................................................................................................. Eremobates

5. Palpal metatarsus, tibia, and femur provided with enlarged spine-like seta, but not robust,
   not aligned in a ventral row, not movable .............................................................. Eremocosta

    Palpal metatarsus, tibia, and femur provided ventrally with robust, obviously movable
   spinelike setae .......................................................................................................... Horribates

6. Fixed cheliceral finger strongly recurved, sigmoid, or S-shaped; mesoventral groove absent;
    both dorsal and ventral flagellum complex setae plumose, moderate sized species ..............
   .....................................................................................................................................Chanbria

    Fixed cheliceral finger style-like or needle-like, straight, curved, or undulate but not
   S-shaped or sigmoid; mesoventral groove present; dorsal flagellum complex setae striate,
   ventral setae striate or plumose ............................................................................................. 7

7. Fixed cheliceral finger with a mesoventral groove that may vary from a distinct crease to
   an elongate hollow or cup; dorsal flagellum complex simple or tubular; small to moderate
   sized species ........................................................................................................... Eremochelis

   Fixed cheliceral finger without a mesoventral groove; dorsal flagellum complex hooked or
   spatulate; tiny to small species ........................................................................... Hemerotrecha

315 - Genus Eremochelis Roewer 1934

Eremochelis Roewer 1934: 570.

Therobates Muma 1951: 85 (synonymized by Muma 1970: 30).


Notes:



Included taxa


  • andreasana group
  • Eremochelis andreasana (Muma 1962)
  • Eremochelis sonorae Muma 1987
  • bilobatus group
  • Eremochelis acrilobatus (Muma 1962)
  • Eremochelis arcus (Muma 1962)
  • Eremochelis bilobatus (Muma 1951)
  • Eremochelis cochiseae Muma 1989
  • Eremochelis cuyamacanus (Muma 1962)
  • Eremochelis flexacus (Muma 1963)
  • Eremochelis giboi Muma 1989
  • Eremochelis kerni Muma 1989
  • Eremochelis lagunensis Vazquez 1991
  • Eremochelis macswaini (Muma 1962)
  • Eremochelis morrisi (Muma 1951)
  • Eremochelis noonani Muma 1989
  • Eremochelis nudus Muma 1963
  • Eremochelis plicatus (Muma 1962)
  • Eremochelis rossi Muma 1987
  • Eremochelis truncus Muma 1987
  • branchi group
  • Eremochelis bechteli Muma 1988
  • Eremochelis bidepressus (Muma 1951)
  • Eremochelis branchi (Muma 1951)
  • Eremochelis coloradensis (Muma 1962)
  • Eremochelis flavus Muma 1989
  • Eremochelis fuscellus Muma 1989
  • Eremochelis gertschi (Muma 1951)
  • Eremochelis insignatus Roewer 1934
  • Eremochelis iviei (Muma 1951)
  • Eremochelis malkini (Muma 1951)
  • Eremochelis medialis (Muma 1951)
  • Eremochelis oregonensis Brookhart & Cushing 2002
  • Eremochelis saltoni Muma 1989
  • Eremochelis tanneri Muma 1989
  • imperialis group
  • Eremochelis imperialis (Muma 1951)
  • Eremochelis kastoni Rowland 1974
  • Eremochelis larreae (Muma 1962)
  • Eremochelis rothi (Muma 1962)
  • Eremochelis undulus Muma 1989
  • striodorsalis group
  • Eremochelis striodorsalis (Muma 1962)

Bibliography: