Subfamily Eremobatinae Kraepelin 1899

Dataminae Kraepelin, 1899a: 240-241.

Eremobatinae Kraepelin, 1901: 116-117; Pocock, 1902c: 59; Roewer, 1934: 555; Muma, 1951: 41; Muma, 1970a 4; Muma, 1976: 14; Muma, 1989:4; Harvey 2003e: 235.

Eremorhaxinae Roewer, 1934: 553 (synonymized by Muma, 1951: 41).

 



Diagnosis:

Eremobatine solpugids possess a single claw on the tip of the first leg. Most solpugids, and indeed, most arachnids, bear two claws. The single claw of the eremobatines, which is flattened and has a broad base, is a unique inherited feature (a synapomorphy) shared by members of this distinctive lineage. Five genera have been recognized: Eremobates (88 species), Eremocosta (13 species), Eremorhax (10 species), Eremothera (2 species) and Horribates (3 species). Members of the genus Horribates possess two claws on the tarsus of the first leg and appear to inappropriately included.




Original description:

Kraepelin 1899 (as Dataminae): 207-208 (key), 240. [see Eremobatidae for full transcription].

 

Subsequent accounts:

Kraepelin 1901 (as Eremobatinae): 28-30 (key), 116, 117. [see Eremobatidae for full transcription].

 

Pocock 1902c: 59:

The recorded Central-American genera are referable to the two following subfamilies:-

a. Mandible with feebly developed stridulating-ridges on the inner side; anterior border of
   carapace straight from side to side. Mandible of male without flagellum, and upper jaw
   untoothed ................................................................................................ EREMOBATINAE.

b. Mandible with strongly developed stridulating-ridges on the inner side; the upper jaw in
   the male toothed and furnished with a flagellum ............................................ DAESIINAE.

Subfam. EREMOBATINAE.

Dataminae, Kraepelin, Jahrb. Hamb. Wiss. Anst. xvi. p. 240 (1899).

Eremobatinae, Kraepelin, Das Tierr., Palpigradi et Solifugae, p. 116 (1901).

This subfamily contains the single genus Eremobates.

 

Roewer, 1934: 553 (key), 555:

553 - 2 Subfamilien der Neuen Welt (Mittelamerika und südwestliches Nordamerika), die sich nach der Gliederung der Beintarsen trennen:

   1 '.  1.-4. Tarsus jeweils 1-gliedrig .................................................. 1. Subfam. Eremorhaxinae

   1 ".  1.-3. Tarsus jeweils 1-gliedrig, 4. Tarsus 3-gliedrig .................. 2. Subfam. Eremobatinae

1. Subfam. Eremorhaxinae novo subfam.

Eremobatidae, deren 1. -4. Tarsus jeweils 1-gliedrig ist (Abb. 319, a, b, g, h). Nur 1

Gattung aus Texas:

1. Gen. Eremorhax novo gen.

555 - 2. Subfam. Eremobatinae s. str.

Lit.: Kraepelin 1899 S. 240 (Dataminae part.) und 1901 S. 116 (Eremobatinae part.).

Eremobatidae, deren 2. und 3. Tarsus jeweils 1-gliedrig und deren 4. Tarsus 3-gliedrig ist
(Abb. 320, a-p).

10 Gattungen aus dem südlichen Nord-Amerika und Mexiko, die sich nach folgender Tabelle ihrer ventralen Bedornung des 2. -4. Tarsus unterscheiden:


Kästner, A. (1933-1935): 193-299.

 

Birula, 1938: 12 (as Eremobathidae [sic]).

 

Muma, 1951: 41:

41 -     KEY TO SUBFAMILIES

    One claw on first leg ........................................................................................ Eremobatinae

   Two claws on first leg ....................................................................................... Therobatinae

SUBFAMILY EREMOBATINAE ROEWER, 1934

Eremobatinae ROEWER, 1934, in Bronn, Klassen und Ordnungen des Tierreichs, vol. 5, div. 4, book 4, P. 553.

Eremorhaxinae ROEWER, 1934, in Bronn, op. cit., vol. 5, div. 4, book 4, p. 553.

Eremobatidae with one claw on the tarsus of the first leg; chelicerae about twice as long as wide. Fixed finger of male chelicera usually style-like, except in Eremothera, new genus, and usually having a ventral or mesoventral groove. Males with or without ctenidia on venter of first post-spiracular segment of abdomen.

KEY TO GENERA

MALES

1.  Fixed finger of chelicera style-like ................................................................................ 2

     Fixed finger sculptured, not style-like .......................................................... Eremothera

2.  Mesoventral groove of fixed finger extending to base of finger, small to moderate-
    sized species ................................................................................................ Eremobates

    Mesoventral groove of fixed finger not extending to base of finger, large species ...
    ........................................................................................................................Eremorhax

 

Muma 1970a: 3,4:

3,4 -   KEY TO SUBFAMILIES AND GENERA OF EREMOBATIDAE
(MALES AND FEMALES)

1. Leg I with 1 claw; large, robust, short legged species ...................... Eremobatinae Roewer-2

    Leg I with 2 claws; small, slender, long legged species ....................... Therobatinae Muma-5

2. Palpus with 2 ventral rows of long, robust, spinelike, erectile and deflectile setae on femur,
   tibia, metatarsus, and tarsus ...................................................................... Horribates Muma

    Palpus with or without spinelike setae; when present, apparently not movable and not on
   metatarsus and tarsus .......................................................................................................... 3

(MALES ONLY)

3. Fixed finger short and dorsally lobate or sculptured ................................ Eremothera Muma

    Fixed finger elongate and need1elike or stylelike ................................................................ 4

4. Mesoventral groove of fixed finger extending to base of finger; apical seta of flagellum-
  complex flattened and plumose ................................................................ Eremobates Banks

    Mesoventral groove of fixed finger not extending to base of finger; apical seta of flagellum-
  complex tubular, at most striate ............................................................... Eremorhax Roewer

5. Dorsal setae of flagellum complex simple and tubular ........................... Eremochelis Roewer

    Dorsal setae of flagellum-complex striate, plumose, spatulate, or otherwise modified .......
   ............................................................................................................................................... 6

6. Fixed finger distinctly sinuate, bent upward and bent or curved downward; dorsal setae
  plumose ......................................................................................................... Chanbria Muma

    Fixed finger straight, weakly curved, undulate, or bent downward at tip; dorsal setae striate,
   spatulate or hooked ............................................................................... Hemerotrecha Banks

SUBFAMILY EREMOBATINAE ROEWER, 1934

Eremorhaxinae Roewer, 1934, p. 558.

Eremobatinae Roewer, 1934, p. 553.

Eremorhaxinae Roe wei, H84, p. 558.

Eremobatinae Roewer, Muma, !951, p. 41

(combination of Roewer's subfamilies).

 

Muma 1976: 14:

14 - Subfamily Eremobatinae Roewer, 1934

This subfamily was erected by Roewer (1934) but Muma (1951) combined Roewer's Eremorhaxinae and Eremobatinae. Four genera of this subfamily are presently recognized, Eremobates Banks, Eremorhax Roewer, Eremothera Muma and Horribates Muma. At present, these genera seem to be restricted to North America.

 

Muma 1989: 3 (key), 4

3, 4 -    Key to Subfamilies and Genera of Eremobatidae

(Males Only)

1. Leg 1 with 1 claw; chelicerae about twice as long as wide; small to large species ...............
   ........................................................................................................Eremobatinae Roewer-2

1'. Leg 1 with 2 claws; chelicerae 2.5–3 times longer than wide; tiny to moderate sized
   species ......................................................................................................... Therobatinae-6

2. Fixed cheliceral finger long, style-like or needle-like; mesoventral groove a crease, slot or
   cup-like structure; moderate-sized to large species .......................................................... 3

2'. Fixed cheliceral finger short, sculptured and flanged; mesoventral groove a trough-like
    slot; moderate sized species ................................................................ Eremothera Muma

3. Mesoventral groove an indistinct hollow or crease that does not extend to the base of
    the fixed finger; movable finger dentition greatly reduced ................................................
   ....................................................................................... Arenotherus Brookhart and Muma

3'. Mesoventral groove a distinct crease, cup, or slot that may or may not extend to base of
   fixed finger; movable finger dentition normal or increased .............................................. 4

4. Mesoventral groove short, not extending to base of fixed finger; apical striate or
   plumose setae of male flagellum complex not obviously modified or flattened .............. 5

4'. Mesoventral groove long, extending to base of fixed finger; apical plumose seta of male
   flagellum complex obviously enlarged or flattened covering part of the mesoventral
   groove..................................................................................................... Eremobates Banks

5'. Palpal metatarsus, tibia, and femur provided with 2 ventral rows of enlarged, robust,
    spine-like, obviously movable setae ....................................................... Horribates Muma

6. Fixed cheliceral finger strongly recurved, sigmoid, or S-shaped; mesoventral groove
    absent; both dorsal and ventral flagellum complex setae plumose, moderate sized
    species........................................................................................................Chanbria Muma

6'. Fixed cheliceral finger style-like or needle-like, straight, curved, or undulate but not
    S-shaped or sigmoid; mesoventral groove present; dorsal flagellum complex setae
    striate, ventral setae striate or plumose ......................................................................... 7

7. Fixed cheliceral finger with a mesoventral groove that varies from one or more indistinct
    creases to an elongate hollow or cup; small to moderate sized species ............................
   ............................................................................................................. Eremochelis Roewer

7'. Fixed cheliceral finger without a mesoventral groove; tiny to small species ......................
    ............................................................................................................ Hemerotrecha Banks

4 -        Subfamily Eremobatinae Roewer

Males and females of this subfamily have one claw on the tarsus of leg 1. This claw is difficult to see on some species of Eremopus Roewer and Horribates Muma but can be observed from an end or tip view at high magnifications. With few exceptions, such as the small Arenotherus mumai (Brookhart), these species are moderate to large in size . Most have chelicerae that are nearly or slightly more than twice as long as wide. Again with a few exceptions, most species are predominately pale to dusty yellow in color with specifically variable purple to purplish brown markings . Except for the description of Horribates Muma (1962), the addition of new species groups in Muma (1970), and the present paper, this subfamily is essentially the same as it was in Muma (1951).

 

Harvey 2003e: 235:

235 -   Subfamily EREMOBATINAE Kraepelin

Dataminae Kraepelin, 1899a: 240-241.

Eremobatinae Kraepelin, 1901: 116-117; Pocock, 1902c: 59; Roewer, 1934: 555; Muma, 1951: 41; Muma, 1970a 4; Muma, 1976: 14; Muma, 1989:4.

Eremorhaxinae Roewer, 1934: 553 (synonymised by Muma, 1951: 41).


Bibliography:


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