Eremocosta spinipalpis (Muma 1970)

Eremocosta striata (Putnam 1883)

Datames spinipalpis Kraepelin, 1899a: 243–244

Kraepelin, 1899b: 378.

Eremobates spinipalpis (Kraepelin): Kraepelin, 1901: 124; Roewer, 1934: 577.

Eremorhax spinipalpis (Kraepelin): Muma, 1970a: 8, fig. 7; Muma, 1976: 15; Polis and McCormick, 1986: 112; Muma, 1987: 19; Vázquez Rojas, 1995: 30; Vázquez Rojas, 1996: 76.

Eremocosta spinipalpis (Kraepelin): Harvey, 2002: 451.

 

Diagnosis:

...


Holotype:

Female holotype from Santa Rosalio, Lower California, México, coll. S. Diguet, Number 33–97 in Muséum National d’Histoire Naturelle, Paris, France.


Original description:

Kraepelin, 1899a: 243–244

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Subsequent accounts:

 

Kraepelin, 1899b: 378.:

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Kraepelin, 1901: 124:

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Roewer, 1934: 577:

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Muma, 1970a: 8, fig. 7:

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Muma 1976: 15:

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Polis and McCormick, 1986: 112:

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Muma, 1987: 19:

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Vázquez Rojas 1995: 30:

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Vázquez Rojas, 1996: 76:

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Harvey 2002: 451:

Recognized Eremopus Roewer 1934 as a junior homonym of the copepod genus Eremopus Brady 1910, resurrected Eremocosta Roewer 1934, transferred Datames striatus Putnam 1883 to Eremocosta.

 

Harvey, 2003:

"Eremocosta striata (Putnam)

Datames striatus Putnam, 1883: 255–257, figs 1, 7–11.

Datames cinerea Putnam, 1883: 260–261, figs 4, 17–31 (synonymised by Muma, 1951: 45).

Eremobates cinereus (Putnam): Banks, 1900: 427 (as Eremobates cinerea [sic]); Banks, 1901b: 589 (as Eremobates cinerea [sic]); Kraepelin, 1901: 124; Comstock, 1913: 37, fig. 33; Roewer, 1934: 577.

Eremorhax striatus (Putnam): Muma, 1951: 45–47, figs 26–31; Muma, 1966c: 201, 207, 212; Muma, 1966d: 256 (as Eremorax [sic] striatus); Kaestner, 1968: 225; Muma, 1970a: 8–9, unnumbered fig. on p. 9; Muma, 1976: 15; Rowland and Reddell, 1976: 7; Cloudsley-Thompson, 1977: 65; Muma, 1985: fig. 3; Muma, 1986: 3–4; Muma, 1987: 3–4, 19; Vázquez Rojas, 1995: 30; Vázquez Rojas, 1996: 76.

Eremopus striatus (Putnam): Muma and Muma, 1988: 10–11, fig. 2; Muma, 1989: 5.

Eremocosta striata (Putnam): Harvey, 2002: 451.

Type localities: of Datames striatus: Camp Grant, Pima County, Arizona, U.S.A. of Datames cinerea: unknown.

Distribution: Mexico, U.S.A. (Arizona, California, Texas)."

 

Brookhart and Brookhart 2006: 304:

"Eremocosta striata (Putnam 1882)

Datames striatus Putnam 1882:255.

Datames cinerea Putnam 1882:260 (synonymized by Muma 1951:45).

Eremobates cinereus (Putnam): Kraepelin 1901: 124.

Erermorhax striatus (Putnam): Muma 1951: 45.

Eremopus striatus (Putnam): Muma & Muma 1988: 10.

Eremocosta striata (Putnam): Harvey 2002: 451.

Type material.Datames striatus: Female holotype from Camp Grant, Arizona, USA (BNHM). Datames cinerea: Type locality of male holotype unknown (ANS).

Recorded specimens.— Several males and females.

Distribution.— USA: Arizona, California. Mexico: Sonora. Biome: Sonoran Desert."

 

Cushing, Graham, Prendini, and Brookhart 2015: 285, 286, 288, 289, Table 2, figs. 1-3:

Cushing et al. included E. striata in a phylogenetic analysis utilizing fragements of the 16S, CO1, H3, and 28S genes.

Figures 2 and 3 depict E. striata as the sister taxon to E. gigasella, with divergence estimated to be in the upper Miocene.

Table 2 reports the locality data for the specimen as USA: Arizona, Santa Cruz Co., east of Amado, 31.71400°, -111.00900° DMNS ZA.21079

The GenBank accession numbers for the specimen are identified as: 16S: KT276625; CO1: KT276707; H3: KT276873; 28S: KT276791.

Bird, Wharton, and Prendini: 8, 97, Fig. 1L:

"An additional process, situated distal to the MM tooth on the gnathal edge of the movable finger is evident in some males, e.g., in Eremochelis imperalis (Muma, 1951), Eremocosta gigas Roewer, 1934, and Eremocosta striata (Putnam, 1883) (Muma, 1951: 46, fig. 26; 46, fig. 32; 95, fig. 174, 175). "

 

Cushing, Channiago and Brookhart:

"Eremocosta striata (Putnam 1883)

  Figs. 1F, 2S–V, 3F, 4E, 5G

     Datames striatus Putnam 1883: 255–257.

     Datames cinerea Putnam 1883: 260–261.

     Eremobates cinereus (Putnam 1883): Kraepelin 1901: 124.

     Eremorhax striatus (Putnam 1883): Muma 1951: 45–47, figs. 26–31.

     Eremopus striatus (Putnam 1883): Muma & Muma 1988: 10–11, fig. 2.

     Eremocosta striata (Putnam 1883): Harvey 2002: 451.

Type material. Datames striatus: Female holotype from Camp Grant, Arizona, USA (BNHM). Datames cinerea: Type locality of male holotype unknown (Academy of Natural Sciences, Philadelphia, Pennsylvania, USA). We did not examine the female holotype.

Specimens examined. Males (8). USA: Arizona, Graham County, 6 miles south of San Carlos, San Carlos Reservoir, N 33.24996°, W 110.44431°, 805 m elev., 5 August 1974, coll. Martin H. Muma (2 males: DMNS ZA.16329); USA: Arizona, Santa Cruz County, E. of Amado along Hopkins Mountain Rd., between the 2–5 Km signs, N 31.714°, W 111.009°, 1011 m elev., 30 July 2008, coll. Joe Warfel (DMNS ZA.21079); USA: Arizona, Santa Cruz County, Hopkins Mountain Rd., toward Whipple Observatory, N 31.67222°, W 110.90528°, 1753 m elev., 28 July 2009, coll. Joe Warfel, on roadway (DMNS ZA.25446); USA: Arizona, Santa Cruz County, Hopkins Mountain Rd., N 31.46139°, W 111.00417°, 1112 m elev., 28 July 2009, coll. Joe Warfel, on roadway at 21:28 hrs (DMNS ZA.25453); USA: Arizona, Pima County, 10 miles E Continental, 3 miles N Madera Canyon, 14–15 July 1989, K. Ribardo (2 males: CAS Casent 9033604).

   Females (2). USA: Arizona, Santa Cruz County, Esplendor Resort, 1069 Camino Caralampi, Rio Rico, N 31.46194°, W 110.99746°, 1063 m elev., July 2009, coll. Lou Perrotti, caught in room (DMNS ZA.25451); MÉXICO: Sonora, 73.9 miles N. of Hermosillo, 5 August 1968, coll. T. Briggs (CAS Casent-9033697).

Diagnosis. The shape of the male chelicera with its slightly rounded fondal notch, a tiny MM, and a flattened ridge-like MD distinguishes this species from all other Eremocosta. Only E. gigasella has a ridge-like structure positionally homologous to the sharper distal ridge that might be a modified MD (see Figs. 2S & O, arrows). Eremocosta gigas has a distinctive MD tooth at that same position on the movable finger (Fig. 2K, arrow). The genital operculum of E. striata is most similar to that of E. calexicensis; however, the curved distal portions of the genital wings are more rounded than those of E. calexicensis (compare Figs. 4E & B).

   Measurements. Males (n = 8). TL 31.0–50.0; CL 9.0–16.0; CH 3.4–5.6; FNL 0.3–2.0; FNH 0.4–0.9; FFH 1.0–1.6; PL 25.0–40.5; PT 7.0–8.0; PMT 1.7–3.0; LI 20.5–31.0; LIV 27.0–43.0; PPL 5.0–7.9; A/CP 4.3–5.6; FNL/FNH 0.5–2.5; FFH/FNH 1.3–2.3; FFH/CH 0.2–0.5.

   Measurements. Females (n = 2). TL 34.5, 50.0; CL 9.0, 17.5; CH 4.0, 5.5; PL 22.0, 35.0; PT 7.5, 7.7; PMT 2.0, 2.9; LI 19.5, 26.0; LIV 34.5, 35.0; PPL 3.9, 4.0; A/CP 3.9, 4.0.

   Description. Coloration. Male. Overall dusky yellow body and appendages; legs dusky violet-brown at tibiafemur joint. Propeltidium reticulate violet-brown with a longitudinal lighter stripe the width of the eye tubercle (Fig. 5G). Abdomen dark dorsally and lighter ventrally.

   Chelicera. Male. VDC apical, oriented prolaterally, narrow, with thin carina (Fig. 1F, arrow). Fixed finger broad at the base narrowing at the apical third of the length in retrolateral view with no median dentition. Movable finger: prominent MP-two tiny MSM (some specimens with just one)-either a very small, but distinct, MM or just a slight ridge at that location (difficult to see in Fig. 2S)-a long ridge-like MD; distinct MPL (Figs. 2S & T). Fondal notch seemingly curved and distinct (Figs. 2S & T). Two tiny RFA; fondal teeth graded I, III, II, IV retrolaterally and I, II, III, IV prolaterally (Fig. 3F).

   Setation. Male. Typical.

   Coloration. Female. Coloration as in the males but sometimes markings are less distinct.

   Chelicera. Female. Fixed finger: large FP-2 small FSM-FM-2 small FSD-smaller FD. Movable finger: large MP-two small MSM-medium MM; smaller but distinct MPL (Figs. 2U & V). Two RFA, proximal tiny; fondal teeth graded as in the male.

   Setation. Female. Typical.

   Genital operculum. Muma’s illustration is accurate (1951, fig. 31, p. 46). Thin anterior arms; relatively large posterior wings (Fig. 4E).

   Distribution. USA Arizona. MÉXICO: Sonora. Biome: Sonoran Desert.

   Discussion. Eremocosta striata is a large species that can be seen by vehicle headlights scurrying across a road (pers. obs, JB). It seems to occupy the Sonoran region east of the Colorado River in Arizona and the state of Sonora, México."


Distribution:

UNITED STATES: Arizona; MEXICO: Sonora.


 


Notes:

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Bibliography: